Dr. David Hoekman

Associate Professor

Email: dhoekman@redeemer.ca

Office: 221C

Programs: Biology, Environmental Science, Environmental Studies


Ph.D. in Ecology, The Graduate School at the University
of Notre Dame, Notre Dame IN, May 2008.

B.S. magna cum laude in Biology and Archaeology, Wheaton College, Wheaton
IL, May 2002.


  • Fundamentals of Biology II (BIO-122)
  • Understanding Our World through the Sciences (SCI-210)
  • Ecology and Evolution (BIO-232)
  • Field Biology (BIO-333)
  • Perspectives on Evolution (BIO-335)
  • Plant Physiology (BIO-343)
  • Senior Seminar I (BIO-381)
  • Senior Seminar II (BIO-382)
  • Senior Seminar III (BIO-383)
  • Senior Seminar IV (BIO-384)


Dr. Hoekman joined Redeemer in Fall, 2020, in the Biology Department. Previously, he was an Associate Professor of Biology at Southern Nazarene University, and the director of the Quetzal Education Research Center.  He received a B.S. in Biology and Archaeology from Wheaton College and a Ph.D. in Ecology from the University of Notre Dame. He studies arthropod food webs in aquatic and terrestrial systems and has taught a variety of courses including Marine Ecology, Insect Ecology, Ornithology, General Ecology, and Tropical Field Research in Costa Rica.

Dr. Hoekman, along with his wife and three sons, enjoys traveling, biking and spending time outside. Prior to coming to Ontario, they have lived and worked in Oklahoma, Colorado, Montana, Wisconsin, Iceland, Indiana and Michigan.

Research Interests

  • Insect ecology
  • Ground-dwelling arthropod communities
  • Aquatic-terrestrial resource pulses
  • Predator-prey relationships
  • Arctic lakes
  • Pitcher plants

Research Funding

“Support for summer student research: ladybeetle biocontrol of aphids in southern Wisconsin” Catalyst Alumni Group research and travel grant, funded in the spring of 2019. ($500)

“Geographic Information System training and research application” Wally Quanstrom Distinguished Faculty Scholar Grant, funded in the spring of 2018 ($10,000)

“Continued research at QERC: arthropod communities along an elevational gradient” Catalyst Alumni Group research and travel grant, funded in the spring of 2018. ($1500)

“Summer research proposal: insect studies at QERC” Catalyst Alumni Group research and travel grant, funded in the spring of 2017. ($1000)

“Developing long-term undergraduate research at QERC: stream macroinvertebrates, bromeliad communities and forest floor arthropods” Southern Nazarene University Faculty Scholarship Support Grant.  Funded in the fall of 2016. ($1000)

“Ecology of Fear: Behavioral Responses of Woodland Deer Mice Peromyscus maniculatus to Direct and Indirect Predator Cues” Catalyst Alumni Group travel grant, funded in the spring of 2016. ($1500)

“Effects of allochthonous aquatic subsidies on terrestrial arthropod food webs: a request for REU supplement” National Science Foundation REU supplement to ecosystems grant, funded in the spring of 2012. ($6850)

“Effects of allochthonous aquatic subsidies on terrestrial arthropod food webs: a request for REU supplement” National Science Foundation REU supplement to ecosystems grant, funded in the spring of 2011. ($7700)

Research Publications

McCary, M. A., M. D. Kasprzak, J. C. Botsch, D. Hoekman, R. D. Jackson, C. Gratton. (2021) Aquatic insect subsidies influence microbial composition and processing of detritus in near-shore subarctic heathland. Oikos, 130(9): 1523-1534. doi: 10.1111/oik.08032

Hoekman, D., M. A. McCary, J. Dreyer, C. Gratton. (2019) Reducing allochthonous resources in a subarctic grassland alters arthropod food webs via predator diet and density. Ecosphere 10(2):e02593. 10.1002/ecs2.2593

Bartrons, M., J. Sardans, D. Hoekman, J. Peñuelas. (2018) Trophic transfer from aquatic to terrestrial ecosystems: a test of the biogeochemical niche hypothesis. Ecosphere 9(7). https://doi.org/10.1002/ecs2.2338

Gratton, C., D. Hoekman, J.M. Dreyer, R.D. Jackson. (2017)  Increased duration of aquatic resource pulse alters community and ecosystem responses in a subarctic plant community. Ecology 98(11): 2860-2872. doi: 10.1002/ecy.1977

Krowiak, A., C.M. Herren, K.C. Webert, Á. Einarsson, D. Hoekman, R.D. Jackson, A.R. Ives. (2017)  Resource gradients and the distribution and flowering of butterwort, a carnivorous plant. Annales Zoologici Fennici 54(1-4): 163-173. doi: http://dx.doi.org/10.5735/086.054.0115

Hoekman, D., K.E. LeVan, C. Gibson, G.E. Ball, R.A. Browne, R.L. Davidson, T.L. Erwin, C.B. Knisley, J.R. LaBonte, J. Lundgren, D.M. Maddison, W. Moore, J. Niemelä, K.A. Ober, D.L. Pearson, J.R. Spence, K. Will and T. Work.  (2017)  Design for Ground Beetle Abundance and Diversity Sampling within the National Ecological Observatory Network. Ecosphere 8(4): e01744. 10.1002/ecs2.1744.

Hoekman, D., Y.P. Springer, C. Gibson, C.M. Barker, R. Barrera, M.S. Blackmore, W.E. Bradshaw, D.H. Foley, H.S. Ginsberg, M. H. Hayden, C. M. Holzapfel, S.A. Juliano, L.D. Kramer, S.L. LaDeau, T.P. Livdahl, C.G. Moore, R.S. Nasci, W.K. Reisen, and H.M. Savage. (2016)  Design for mosquito abundance, diversity, and phenology sampling within the National Ecological Observatory Network. Ecosphere 7(5):e01320. 10.1002/ecs2.1320.

Thorpe, A.S., D. Barnett, S. Elmendorf, E. Hinckley, D. Hoekman, K. Jones, K. LeVan, C. Meier, L. Stanish, K. Thibault. (2016)  Introduction to the sampling designs of the National Ecological Observatory Network Terrestrial Observation System. Ecosphere 7(12): doi 10.1002/ecs21627.

Springer, Y.P., D. Hoekman, P.T.J. Johnson, P.A. Duffy, R.A. Hufft, D.T. Barnett, B.F. Allan, B.R. Amman, C.M. Barker, R. Barrera, C.B. Beard, L. Beati, M. Begon, M.S. Blackmore, W.E. Bradshaw, D. Brisson, C.H. Calisher, J.E. Childs, M.A. Diuk-Wasser, R.J. Douglass, R.J. Eisen, D.H. Foley, J.E. Foley, H.D. Gaff, S.L. Gardner, H.S. Ginsberg, G.E. Glass, S.A. Hamer, M.H. Hayden, B. Hjelle, C.M. Holzapfel, S.A. Juliano, L.D. Kramer, A.J. Kuenzi, S.L. LaDeau, T.P. Livdahl, J.N. Mills, C.G. Moore, S. Morand, R.S. Nasci, N.H. Ogden, R.S. Ostfeld, R.R. Parmenter, J. Piesman, W.K. Reisen, H.M. Savage, D.E. Sonenshine , A. Swei, M.J. Yabsley. (2016) Continental-scale surveillance of infectious agents: Tick-, mosquito-, and rodent-borne parasite sampling designs for NEON. Ecosphere 7(5):e01271. 10.1002/ecs2.1271.

Dreyer, J.M., D. Hoekman, C. Gratton. (2015) Positive indirect effect of aquatic insects on terrestrial prey is not offset by increased predator density.  Ecological Entomology, doi: 10.1111/een.12272.

Sabo, J.L., D. Hoekman. (2015) Chapter 6: Dynamic systems of exchange link trophic dynamics in freshwater and terrestrial food webs. Editors: Kim La Pierre and Torrie Hanley.  Trophic Ecology: Bottom-Up and Top-Down Interactions Across Aquatic and Terrestrial Systems.  Cambridge University Press.

Dreyer, J.M., P.A. Townsend, J.C. Hook III, D. Hoekman, M.J. Vander Zanden, C. Gratton. (2015) Quantifying aquatic insect deposition from lake to land. Ecology 96(2):499-509, doi.org/10.1890/14-0704.1.

Bultman, H., D. Hoekman, J. Dreyer, C. Gratton. (2014) Terrestrial deposition of aquatic insects increases plant quality for insect herbivores and herbivore density.  Ecological Entomology 39(4):419-426, doi: 10.1111/een.12118.

Davies, N., D. Field, L. Amaral-Zettler et al. (69 authors including D. Hoekman and other NEON scientists). (2014) The founding charter of the Genomic Observatories Network. GigaScience 3(2), doi:10.1186/2047-217X-3-2.

Hoekman, D., M. Bartrons, C. Gratton. (2012) Ecosystem linkages revealed by experimental lake-derived isotope signal in heathland food webs. Oecologia 170(3) 735-743.

Choate, D.M, C.M. Prather, M.J. Michel, A.K. Baldridge, M.A. Barnes, D. Hoekman, C.J. Patrick, J. Rüegg and T.A. Crowl. (2012) Integrating Theoretical Components: a Graphical Model for Graduate Students and Researchers. Bioscience 62(6): 594-602.

Dreyer, J.M., D. Hoekman, C. Gratton. (2012) Lake derived aquatic insects increase abundance of terrestrial arthropods, Oikos 121(2): 252-258.

Hoekman, D., J.M. Dreyer, R.D. Jackson, P.A. Townsend, C. Gratton. (2011) Lake to land subsidies: experimental addition of aquatic insects increases terrestrial arthropod densities. Ecology 92(11): 2063-2072.

Hoekman, D. (2011) Sarracenia purpurea pitcher communities at disparate latitudes: a field study of the relative importance of top-down and bottom-up forces in food webs. Oecologia 165(4): 1073-1082.

Hoekman, D. (2010) The effect of temperature on the relative importance of top-down and bottom-up effects. Ecology 91(10): 2819-2825.

Hoekman, D., R. Winston, and N. Mitchell. (2009) Top-down and bottom-up effects of a processing detritivore. Journal of the North American Benthological Society 28(3): 552-559.

Hoekman, D., C. Terhorst, A. Bauer, S. Braun, P. Gignac, R. Hopkins, S. Joshi, K. Laskis, N. Sanscrainte, J. Travis, and T.E. Miller. (2007) Oviposition decreased in response to enriched water:  a field study of the pitcher-plant mosquito, Wyeomyia smithii. Ecological Entomology 32: 92-96.

Hoekman, D. (2007) Top-down and bottom-up regulation in a detritus-based aquatic food web: a repeated field experiment using the pitcher plant (Sarracenia purpurea) inquiline community. American Midland Naturalist 157: 52-62.

Off-Campus Teaching

Au Sable Institute of Environmental Studies (2016-current): BIOL 365 Insect Ecology of Streams, Forests, and Fields. (lecture & lab)

Wheaton College Science Station (2015, 2016, 2021): BIOL 243 Processes of Life: Ecology and Evolution (lecture & lab)